AP Psych: Hemispheric Specialization Definition + Examples


AP Psych: Hemispheric Specialization Definition + Examples

Cerebral lateralization refers to the concept that the two hemispheres of the brain are not functionally equivalent. Instead, each hemisphere exhibits specialization for particular cognitive and behavioral processes. For instance, the left hemisphere is typically dominant for language processing, controlling speech, grammar, and comprehension in most individuals. Conversely, the right hemisphere often demonstrates superiority in visuospatial tasks, facial recognition, and emotional processing. This division of labor allows for more efficient cognitive function. The specific functions associated with each side of the brain can vary slightly among individuals.

Understanding the specific functions of each side of the brain is crucial for diagnosing and treating neurological disorders. For example, damage to the left hemisphere can result in aphasia, an impairment of language abilities. Conversely, damage to the right hemisphere may lead to deficits in spatial awareness or the ability to recognize familiar faces. Awareness of this functional segregation has evolved through historical research, including studies of split-brain patients who had their corpus callosum severed, revealing distinct hemispheric capabilities. These studies highlight the independent processing capacities of each hemisphere when communication between them is disrupted.

The functional differences between the two sides of the brain are significant in many areas of psychology. These concepts are vital in studying perception, memory, language, and other core cognitive functions. This lateralization also plays a role in our everyday actions and behavior.

1. Lateralization of function

Lateralization of function is a foundational concept directly related to the understanding of hemispheric specialization within the field of psychology. It posits that specific cognitive and behavioral functions are primarily controlled or processed by one hemisphere of the brain over the other. This principle forms the basis for defining how the brain divides labor, leading to functional differences between the left and right hemispheres.

  • Language Processing and Left Hemisphere Dominance

    The left hemisphere is predominantly responsible for language functions in most individuals. This lateralization manifests as proficiency in speech production, grammar comprehension, and vocabulary access. Damage to specific areas within the left hemisphere, such as Broca’s area or Wernicke’s area, can result in various forms of aphasia, directly demonstrating the critical role of this hemisphere in language. This illustrates a clear instance of functional segregation and its implications for cognitive abilities.

  • Visuospatial Skills and Right Hemisphere Specialization

    The right hemisphere is typically associated with enhanced visuospatial processing. This includes abilities such as facial recognition, spatial orientation, and the perception of depth. Individuals with damage to the right hemisphere may exhibit difficulties in navigating familiar environments or recognizing faces, underscoring the hemisphere’s crucial role in these domains. This highlights another core aspect of the distinct processing strengths of each hemisphere.

  • Emotional Processing and Hemispheric Asymmetry

    While both hemispheres contribute to emotional processing, the right hemisphere plays a significant role in recognizing and interpreting emotional cues, particularly negative emotions. Research suggests that the right hemisphere is more responsive to emotional stimuli, and damage to this hemisphere can impair the ability to perceive and understand emotional expressions in others. This contributes to the overall understanding of how emotional intelligence can be linked to specific brain regions.

  • Impact of Split-Brain Studies on Understanding Lateralization

    Split-brain studies, where the corpus callosum connecting the two hemispheres is severed, provide critical insights into hemispheric specialization. These studies reveal that each hemisphere can function independently, exhibiting unique cognitive capacities when communication between them is limited. Split-brain patients can, for instance, identify objects presented to the right hemisphere (left visual field) but struggle to verbally name them due to language dominance in the left hemisphere. These studies offer compelling evidence for the distinct processing capabilities of each hemisphere and their separate contributions to overall cognition.

These facets illustrate the depth and complexity of lateralization of function and its importance in elucidating the concept of hemispheric specialization. By examining language, visuospatial skills, emotional processing, and the insights gained from split-brain research, a more complete picture emerges of how the brain divides labor and optimizes cognitive function through distinct hemispheric specializations.

2. Left hemisphere dominance

Left hemisphere dominance, a central aspect of hemispheric specialization, refers to the preferential control of particular cognitive functions by the left side of the brain. This dominance is most prominently observed in language processing for a majority of individuals. Language centers such as Broca’s area, responsible for speech production, and Wernicke’s area, crucial for language comprehension, are typically located within the left hemisphere. Consequently, damage to these areas, often resulting from stroke, can lead to various forms of aphasia, impairing the ability to speak or understand language. This direct link between the left hemisphere and language function underscores its dominant role in this specific cognitive domain.

Beyond language, the left hemisphere also exhibits dominance in areas such as analytical reasoning, sequential processing, and the control of fine motor movements, particularly on the right side of the body. Its structured approach to information processing allows for the efficient execution of tasks requiring logical analysis and detailed sequencing. For instance, mathematical calculations and the step-by-step breakdown of complex problems often rely heavily on the left hemisphere’s capabilities. Similarly, the precise movements required for writing are typically controlled by the left hemisphere, influencing right-handedness in most individuals.

The understanding of left hemisphere dominance is crucial within the framework of hemispheric specialization. It illustrates that the brain is not a homogenous organ where all functions are equally distributed. Instead, it demonstrates a division of labor, with the left hemisphere taking precedence in certain cognitive and motor domains. This specialization enhances overall cognitive efficiency by allowing each hemisphere to develop specialized processing strategies. Moreover, it offers practical applications in diagnosing and treating neurological conditions, informing rehabilitation strategies for individuals with brain damage, and providing insights into cognitive development and individual differences in abilities.

3. Right hemisphere specialization

Right hemisphere specialization represents a critical component of the broader understanding of hemispheric specialization within the brain. It highlights the distinct functions and cognitive processes primarily managed by the right cerebral hemisphere, contributing to a more complete understanding of how the brain divides labor. This is crucial to the study of psychology.

  • Visuospatial Processing

    The right hemisphere exhibits superiority in processing spatial information, including depth perception, spatial orientation, and the ability to mentally manipulate objects in space. This specialization is evident in tasks such as map reading, solving puzzles, and navigating through complex environments. For instance, individuals with damage to the right hemisphere may exhibit difficulties in recognizing familiar landmarks or in judging distances, directly illustrating its role in spatial cognition.

  • Facial Recognition

    Facial recognition is another key area of right hemisphere specialization. The right hemisphere plays a significant role in processing and identifying faces, including both familiar and unfamiliar ones. Damage to specific regions within the right hemisphere, such as the fusiform face area, can result in prosopagnosia, or face blindness, where individuals lose the ability to recognize faces, even those of close family members. This demonstrates the critical involvement of the right hemisphere in this complex perceptual task.

  • Emotional Processing and Expression

    The right hemisphere contributes to emotional processing and expression. It is particularly involved in recognizing and interpreting nonverbal cues, such as facial expressions and tone of voice, which convey emotional information. This specialization allows individuals to accurately perceive and respond to the emotions of others. Right hemisphere damage may lead to difficulties in understanding emotional signals, impacting social interactions and interpersonal relationships.

  • Artistic and Musical Abilities

    Artistic and musical abilities often rely on the right hemisphere’s strengths in holistic processing and pattern recognition. While both hemispheres contribute to creative endeavors, the right hemisphere is particularly involved in appreciating and creating visual art and music. Its capacity to perceive patterns, spatial relationships, and emotional nuances enhances artistic expression and musical interpretation, contributing to the aesthetic experiences.

These specialized functions of the right hemisphere, encompassing visuospatial processing, facial recognition, emotional processing, and artistic abilities, underscore the critical role it plays in the overall cognitive landscape. Right hemisphere specialization not only complements the functions of the left hemisphere but also contributes uniquely to higher-order cognitive processes, thus completing a comprehensive understanding of cerebral lateralization and behavior.

4. Split-brain studies

Split-brain studies provide pivotal empirical evidence supporting the concept of hemispheric specialization. These studies, conducted on individuals who have undergone a corpus callosotomy, offer unique insights into the independent functions of the left and right cerebral hemispheres. By examining the cognitive and perceptual abilities of these patients, researchers have been able to delineate the specific roles each hemisphere plays in various tasks, thus directly contributing to the understanding of hemispheric specialization.

  • Visual Field Presentation and Hemispheric Access

    In split-brain studies, stimuli presented to one visual field are primarily processed by the contralateral hemisphere. For example, an image flashed briefly to the right visual field is initially processed by the left hemisphere, while an image presented to the left visual field is processed by the right hemisphere. This controlled presentation allows researchers to assess the capabilities of each hemisphere in isolation. If a split-brain patient is shown an object in the left visual field, they may be unable to verbally name the object due to the language centers location in the left hemisphere, even though their right hemisphere recognizes it. This demonstrates the left hemisphere’s dominance in language and the right hemisphere’s capacity for visual recognition.

  • Conflicting Hemispheric Responses

    Split-brain patients often exhibit conflicting responses when tasks require the integration of information from both hemispheres. If each hemisphere receives conflicting information, the patient may perform actions driven by one hemisphere while simultaneously expressing a different intention verbally, which originates from the other hemisphere. This is exemplified when the right hemisphere instructs the left hand to select an object related to an image presented to the left visual field, while the left hemisphere (controlling speech) provides a different, unrelated verbal response. Such observations underscore the importance of the corpus callosum in normal interhemispheric communication and the independent processing capabilities of each hemisphere.

  • Emotional Processing Asymmetries

    Research using split-brain patients has also revealed asymmetries in emotional processing. The right hemisphere appears to play a more significant role in the recognition and expression of emotions. Studies have shown that the right hemisphere is better at identifying emotional facial expressions than the left. Additionally, when presented with emotionally charged images in the left visual field, split-brain patients exhibit stronger emotional responses, indicating the right hemisphere’s dominance in emotional processing. This evidence further supports the notion of hemispheric specialization in the emotional domain.

  • Limitations and Interpretations

    It is crucial to acknowledge limitations in generalizing from split-brain studies. Split-brain patients represent a unique population with specific neurological alterations that may not be representative of the general population. However, the consistent patterns observed in these studies provide valuable insights into the functional organization of the brain. The findings from split-brain research have been instrumental in shaping theories of hemispheric specialization and have spurred further investigation into the neural basis of cognition and behavior. Interpretations of these studies must consider the compensatory mechanisms that may develop in split-brain patients over time, as well as the variability in cognitive abilities among individuals.

The consistent patterns observed in split-brain studies, relating visual field presentation, hemispheric access, conflicting responses, and emotional processing, provide empirical support to the principles of functional differences between the two sides of the brain.

5. Language processing (left)

Language processing, predominantly localized in the left cerebral hemisphere for the majority of individuals, constitutes a critical component of hemispheric specialization. This lateralization manifests through specialized neural structures and functions within the left hemisphere dedicated to decoding and producing linguistic information. Broca’s area, situated in the frontal lobe, governs speech production, enabling the formulation and articulation of words and sentences. Wernicke’s area, located in the temporal lobe, facilitates language comprehension, allowing the interpretation of spoken and written language. The left hemisphere dominance in these language functions exemplifies the brain’s inherent division of labor and functional segregation.

The impact of left hemisphere specialization for language becomes evident through clinical observations of individuals with damage to these critical language areas. For example, a stroke affecting Broca’s area can result in expressive aphasia, characterized by impaired speech production despite intact comprehension. Conversely, damage to Wernicke’s area can lead to receptive aphasia, where comprehension is severely compromised, although speech may remain fluent but nonsensical. These clinical scenarios underscore the practical significance of understanding hemispheric specialization in diagnosing and treating language-related neurological disorders. Rehabilitation strategies often target the specific language deficits resulting from left hemisphere damage, aiming to restore or compensate for the impaired functions. Furthermore, this knowledge informs the development of educational approaches designed to enhance language acquisition and processing skills.

The intricate relationship between left hemisphere language processing and hemispheric specialization reveals the brain’s remarkable capacity for functional organization. This lateralization not only enhances cognitive efficiency but also allows for complex cognitive tasks, such as language, to be processed with greater precision and speed. While the right hemisphere contributes to certain aspects of language, such as prosody and emotional tone, the left hemisphere remains the dominant force in language processing for most individuals. Acknowledging this asymmetry is crucial for a comprehensive understanding of human cognition and its neural underpinnings. Challenges remain in fully elucidating the complexities of language processing, including the interaction between the two hemispheres and the individual variability in lateralization patterns. However, continued research holds the potential to deepen our understanding and improve diagnostic and therapeutic interventions.

6. Visuospatial skills (right)

The specialization of the right cerebral hemisphere for visuospatial skills represents a cornerstone of hemispheric specialization, illustrating the functional division of labor within the brain. This specialization refers to the enhanced ability of the right hemisphere to process visual and spatial information, enabling individuals to perform tasks involving spatial orientation, object manipulation, and pattern recognition.

  • Spatial Orientation and Navigation

    The right hemisphere is critically involved in spatial orientation, enabling individuals to navigate environments and maintain a sense of direction. This skill is essential for everyday tasks such as finding one’s way through a building, driving a car, or using a map. Damage to the right hemisphere can impair spatial orientation, leading to difficulties in navigation and an increased risk of getting lost. This highlights the right hemisphere’s contribution to cognitive maps and spatial awareness.

  • Mental Rotation and Object Manipulation

    The right hemisphere demonstrates proficiency in mental rotation, the ability to mentally rotate two-dimensional or three-dimensional objects in space. This skill is vital for tasks such as assembling furniture, visualizing the components of a machine, or performing engineering design. Right hemisphere lesions can impair mental rotation abilities, affecting performance in technical fields and everyday problem-solving. This function underscores the capacity for abstract spatial reasoning localized in the right side of the brain.

  • Facial Recognition and Spatial Configuration

    While facial recognition involves multiple brain areas, the right hemisphere plays a significant role in processing the spatial configuration of facial features. This aspect of facial processing enables individuals to recognize familiar faces and differentiate between subtle variations in facial expressions. Damage to specific regions within the right hemisphere can lead to prosopagnosia, the inability to recognize faces, demonstrating the right hemisphere’s contribution to holistic face perception and spatial encoding of facial features.

  • Artistic Abilities and Visual Creativity

    The right hemisphere’s specialization extends to artistic abilities and visual creativity. Its proficiency in processing spatial relationships, patterns, and visual details contributes to the creation and appreciation of visual art. Artists often rely on the right hemisphere’s intuitive processing to generate novel visual representations and to perceive aesthetic qualities in the environment. Right hemisphere activation is associated with creativity and the ability to think visually, further illustrating its dominance in visuospatial domains.

These facets demonstrate the distinct contributions of the right hemisphere to visuospatial skills, which is a core component of hemispheric specialization. Understanding these specialized functions is crucial for diagnosing and treating neurological disorders affecting spatial cognition, as well as for appreciating the complex interplay between the brain’s hemispheres in shaping cognitive abilities.

7. Emotional processing

Emotional processing is significantly lateralized within the brain, forming a key aspect of hemispheric specialization. While both hemispheres contribute to emotional experience, they do so in distinct and often complementary ways. This division of labor is crucial for a nuanced understanding of emotional responses and their neurological underpinnings.

  • Right Hemisphere Dominance for Negative Emotions

    The right hemisphere exhibits a dominant role in processing negative emotions, such as fear, sadness, and anger. This specialization involves both the recognition of negative emotional cues in others and the experience of these emotions. For example, studies show that individuals with damage to the right hemisphere may have difficulty recognizing negative emotional expressions in faces or voices. Furthermore, increased activity in the right hemisphere has been observed during the experience of negative emotions. This lateralization has implications for understanding emotional disorders, where an imbalance in hemispheric activity may contribute to heightened or blunted emotional responses.

  • Left Hemisphere Involvement in Positive Emotions

    While the right hemisphere is more associated with negative emotions, the left hemisphere has been linked to the processing of positive emotions, such as happiness and joy. This lateralization is not as pronounced as the right hemisphere’s dominance for negative emotions, but it is nonetheless significant. Research suggests that increased activity in the left hemisphere is associated with feelings of well-being and optimism. Moreover, damage to the left hemisphere can sometimes result in depressive symptoms, indicating its role in maintaining positive emotional states. Understanding this balance is crucial for comprehending the neural mechanisms underlying mood regulation.

  • Amygdala Lateralization in Emotional Memory

    The amygdala, a key structure in emotional processing, also exhibits lateralization. The left amygdala is often associated with the conscious recollection of emotional events, while the right amygdala is more involved in the unconscious, automatic responses to emotional stimuli. For instance, studies have shown that the left amygdala is activated when individuals deliberately recall emotional memories, whereas the right amygdala responds rapidly and automatically to threatening stimuli. This division highlights the complexity of emotional memory and the different roles of the hemispheres in processing emotional information at both conscious and unconscious levels.

  • Hemispheric Interaction and Emotional Regulation

    Effective emotional regulation requires the interaction and coordination of both hemispheres. The prefrontal cortex, which plays a critical role in regulating emotions, receives input from both hemispheres and exerts control over subcortical structures involved in emotional processing, such as the amygdala. While the right hemisphere may generate emotional responses, the left hemisphere can modulate these responses through cognitive appraisal and emotional suppression strategies. This interplay between the hemispheres is essential for maintaining emotional stability and adapting to changing environmental demands. Disruptions in this communication can lead to emotional dysregulation, contributing to conditions such as anxiety and depression.

The multifaceted nature of emotional processing within the context of hemispheric specialization underscores the complexity of brain function. While certain emotions or aspects of emotional processing may be preferentially associated with one hemisphere, the effective integration and regulation of emotions depend on the coordinated activity of both hemispheres. This division of labor allows for a nuanced and adaptive emotional repertoire, essential for navigating the social and emotional landscape. Future research should continue to explore the dynamic interactions between the hemispheres in emotional processing, with the goal of improving our understanding of emotional disorders and developing more effective treatments.

8. Corpus callosum

The corpus callosum, the largest white matter structure in the brain, serves as the primary commissure connecting the left and right cerebral hemispheres. Its role is intrinsically linked to the functional division observed in hemispheric specialization. The corpus callosum facilitates interhemispheric communication, enabling the transfer of sensory, motor, and cognitive information between the two sides of the brain. Disruption of this communication, as seen in individuals who have undergone a corpus callosotomy (split-brain procedure), can reveal the independent processing capabilities of each hemisphere. This directly supports the concept that each hemisphere possesses unique specializations. For example, while the left hemisphere may be dominant for language, the right hemisphere excels in visuospatial tasks. The corpus callosum normally ensures seamless integration of these specialized functions, allowing for a unified conscious experience. Without it, the distinct functions of each hemisphere become more apparent, leading to observable differences in behavior and cognitive processing.

The significance of the corpus callosum in understanding functional differences is further highlighted by studies examining individuals with agenesis of the corpus callosum (ACC), a congenital condition in which the corpus callosum is partially or completely absent. These individuals often exhibit compensatory mechanisms that allow them to function relatively normally, but detailed neuropsychological testing may reveal subtle deficits in interhemispheric transfer and integration. Examining these deficits provides valuable insights into the specific contributions of the corpus callosum to cognitive functions such as bimanual coordination, attention, and higher-level reasoning. The study of ACC also underscores the brain’s plasticity and its ability to adapt to structural anomalies, potentially influencing the manifestation of hemispheric specialization.

In summary, the corpus callosum is fundamental for understanding the interplay between the two hemispheres, and how they perform different tasks. Its presence enables communication, facilitating integrated cognitive processing and a unified consciousness. Examination of individuals with disruptions to the corpus callosum, whether through surgical intervention or congenital absence, provides critical empirical evidence supporting the theory of distinct functional localization. Continued research into the corpus callosum’s role in hemispheric communication promises to refine our understanding of the brain’s organization and function, offering insights into neurological conditions and informing targeted therapeutic interventions.

Frequently Asked Questions About Hemispheric Specialization

The following questions address common inquiries regarding functional lateralization and its implications for behavior and cognition.

Question 1: Is the concept of “brain dominance” an accurate reflection of cerebral lateralization?

The term “brain dominance” is an oversimplification. While each hemisphere exhibits specialization, neither side is entirely dominant. Most cognitive processes require the cooperation of both hemispheres. Furthermore, the degree of specialization can vary among individuals.

Question 2: Does hemispheric specialization imply that individuals are either “left-brained” or “right-brained”?

The notion of distinct “left-brained” or “right-brained” personalities is a popular misconception. While the hemispheres exhibit preferences for certain functions, cognitive processes typically involve the integrated activity of both hemispheres. Personality traits are far more complex than simple hemispheric dominance.

Question 3: How do split-brain studies contribute to the understanding of hemispheric specialization?

Split-brain studies, involving individuals who have undergone a corpus callosotomy, provide unique insights into the independent capabilities of each hemisphere. By presenting stimuli to one visual field, researchers can assess the specialized functions of the contralateral hemisphere in isolation, revealing its distinct processing abilities.

Question 4: What are the clinical implications of understanding hemispheric specialization?

Understanding functional lateralization is crucial for diagnosing and treating neurological disorders. Damage to specific areas within one hemisphere can result in predictable cognitive or behavioral deficits, allowing for targeted interventions and rehabilitation strategies. Knowledge of hemispheric specialization can inform the diagnosis and treatment of conditions such as aphasia, spatial neglect, and prosopagnosia.

Question 5: Can the functions associated with one hemisphere be transferred to the other following brain damage?

The brain exhibits plasticity, allowing for some degree of functional reorganization following injury. In certain cases, the undamaged hemisphere may assume some of the functions previously performed by the damaged hemisphere. However, the extent of this transfer depends on factors such as the age of the individual, the severity of the injury, and the specific functions involved.

Question 6: Does hemispheric specialization change throughout the lifespan?

Hemispheric specialization develops throughout childhood and adolescence, with certain functions becoming more lateralized over time. While the degree of lateralization may remain relatively stable in adulthood, aging can sometimes lead to a decline in hemispheric asymmetry. However, the precise nature and extent of these changes are complex and vary among individuals.

Hemispheric specialization is a nuanced concept with significant implications for cognitive function. A comprehensive understanding of lateralization requires a departure from simplistic notions of “brain dominance” and a recognition of the complex interplay between the two cerebral hemispheres.

The next section explores the future directions of research and the evolving understanding of hemispheric specialization.

Tips for Mastering Hemispheric Specialization in AP Psychology

This section provides advice for mastering the concept of hemispheric specialization, a vital area of study within AP Psychology. Successfully grappling with the definition requires focused effort and a comprehensive approach.

Tip 1: Define the Core Concept Precisely: It’s important to have a well-defined concept of what specialization means in the context of the brain. Note specific functions more dominant on one side than the other. Avoid generalizations; focus on researched findings.

Tip 2: Memorize Key Functional Asymmetries: Commit to memory the most important functions of the brain. Ensure the left hemisphere is associated with language processing (Broca’s and Wernicke’s areas), and the right hemisphere is associated with visuospatial skills and facial recognition.

Tip 3: Master Split-Brain Studies: Use the results of split-brain studies as the primary data to explain the differences between each side. Know how experiments with split-brain individuals shed light on independent hemispheric processing when the corpus callosum is severed.

Tip 4: Acknowledge the Limits of Lateralization: Be cognizant that while there are differences, these differences aren’t set in stone and certain functions need both sides to perform. Recognize that in real life, functions depend on both sides of the brain that communicate.

Tip 5: Study Lateralization of Sensory and Motor Functions: The left brain controls right side body movements while the right brain controls left side body movements. Do not assume that sensory processing works the same way as motor.

Tip 6: Understand the Corpus Callosum: It is a vital white matter structure that connect two hemispheres. Ensure the effects of the corpus callosum are understood by doing various actions simultaneously from both sides.

Tip 7: Connect to Clinical Applications: Know how to connect specific functions to their roles on the brain. Link this concept with neurological conditions, such as aphasia or prosopagnosia, to understand the practical implications of functional differences.

Mastering the hemispheric specialization definition in AP Psychology necessitates a balance between memorization and critical thinking. By focusing on the core definitions, key functional asymmetries, split-brain research, and the role of the corpus callosum, successful mastery of this important concept can be achieved.

The next section offers a comprehensive conclusion that reinforces the importance of the hemispheric specialization definition.

Hemispheric Specialization in Psychology

This exploration of the hemispheric specialization ap psychology definition has illuminated the functional divisions within the brain’s left and right cerebral hemispheres. The examination has underscored that cognitive and behavioral functions are not uniformly distributed, but rather exhibit preferential localization to one hemisphere or the other. Language processing, particularly speech production and comprehension, is predominantly associated with the left hemisphere. Conversely, the right hemisphere demonstrates superiority in visuospatial skills, facial recognition, and certain aspects of emotional processing. The study of split-brain patients, who have undergone a corpus callosotomy, has provided invaluable insights into the independent processing capabilities of each hemisphere, while acknowledging that these patients represent a unique population with potential compensatory mechanisms.

Understanding the hemispheric specialization ap psychology definition is critical for appreciating the complexity of human cognition and behavior. As research progresses, the intricacies of interhemispheric interaction and the dynamic interplay between specialized functions will be further elucidated. Continued investigation into the neural basis of functional differences is essential for advancing diagnostic and therapeutic interventions for neurological disorders. It is critical to look at these studies in order to expand cognitive and neuroscience research.